Phylogenetic placement of diversification rate shifts and pharyngeal jaw modifications. Tip clade richness follows names with warmer colored tip triangles indicate subclades with greater species richness. Numbered branches indicate position of two diversification rate increases. Origin of labroid pharyngeal jaw apparatus (PJM) and parrotfish pharyngeal mill (PM) indicated by black rectangles. Tree backbone is taken from Figure 1. Species richness and taxonomic membership of major subclades given in Table 8.