Extent of variation in phenotypic traits analyzed by individual nested ANOVAs for 14 morphological characters. Colonies are nested within habitats. Black and white bars represent samples in shallow and deep areas, respectively. Values are shown as mean value ± SE (n = 300 measurements per location for micromorphological characters and 15 per location for macromorphological characters). Lowercase letters indicate treatments that are statistically different by post hoc analysis. The location codes are as in Table 6.