From: The evolution of competition and policing: opposing selection within and among groups
Half sib
Full sib
Among-family variance in policing (σ2a_AF)
tua a 2/2
tua a 2
Mean fitness (W)
(1 - c a ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacuWGHbqygaqeaaaa@2E0F@ )(1 - z(1 - a ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacuWGHbqygaqeaaaa@2E0F@ )) - czσ2 a_AF_HS
(1 - c a ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacuWGHbqygaqeaaaa@2E0F@ )(1 - z(1 - a ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacuWGHbqygaqeaaaa@2E0F@ )) - czσ2 a_AF_FS
Δu within groups (Δuw)
− a a c t u 4 w ¯ H S ( 3 − z ( 3 − ( 2 a ¯ + a 2 H S ) ) ) MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaadaWcaaqaaiabgkHiTiabdggaHnaaBaaaleaacqWGHbqyaeqaaOGaem4yamMaemiDaqNaemyDauhabaGaeGinaqJafm4DaCNbaebadaWgaaWcbaGaemisaGKaem4uamfabeaaaaGcdaqadaqaaiabiodaZiabgkHiTiabdQha6naabmaabaGaeG4mamJaeyOeI0YaaeWaaeaacqaIYaGmcuWGHbqygaqeaiabgUcaRiabdggaHnaaBaaaleaacqaIYaGmcqWGibascqWGtbWuaeqaaaGccaGLOaGaayzkaaaacaGLOaGaayzkaaaacaGLOaGaayzkaaaaaa@4B77@
− a a c t u 2 w ¯ F S ( 1 − z ( 1 − ( a ¯ + a 4 F S ) / 2 ) ) MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaadaWcaaqaaiabgkHiTiabdggaHnaaBaaaleaacqWGHbqyaeqaaOGaem4yamMaemiDaqNaemyDauhabaGaeGOmaiJafm4DaCNbaebadaWgaaWcbaGaemOrayKaem4uamfabeaaaaGcdaqadaqaaiabigdaXiabgkHiTiabdQha6naabmaabaGaeGymaeJaeyOeI0YaaeWaaeaacuWGHbqygaqeaiabgUcaRiabdggaHnaaBaaaleaacqaI0aancqWGgbGrcqWGtbWuaeqaaaGccaGLOaGaayzkaaGaei4la8IaeGOmaidacaGLOaGaayzkaaaacaGLOaGaayzkaaaaaa@4C4D@
Δu between groups (Δub)
a a t u 4 w ¯ H S ( − c + z ( 1 + c ( 1 − ( a ¯ + a 2 H S ) ) ) ) MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaadaWcaaqaaiabdggaHnaaBaaaleaacqWGHbqyaeqaaOGaemiDaqNaemyDauhabaGaeGinaqJafm4DaCNbaebadaWgaaWcbaGaemisaGKaem4uamfabeaaaaGcdaqadaqaaiabgkHiTiabdogaJjabgUcaRiabdQha6naabmaabaGaeGymaeJaey4kaSIaem4yam2aaeWaaeaacqaIXaqmcqGHsisldaqadaqaaiqbdggaHzaaraGaey4kaSIaemyyae2aaSbaaSqaaiabikdaYiabdIeaijabdofatbqabaaakiaawIcacaGLPaaaaiaawIcacaGLPaaaaiaawIcacaGLPaaaaiaawIcacaGLPaaaaaa@4E2C@
a a t u 2 w ¯ F S ( − c + z ( 1 + c ( 1 − ( a ¯ + a 2 H S ) ) ) ) MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaadaWcaaqaaiabdggaHjabdggaHjabdsha0jabdwha1bqaaiabikdaYiqbdEha3zaaraWaaSbaaSqaaiabdAeagjabdofatbqabaaaaOWaaeWaaeaacqGHsislcqWGJbWycqGHRaWkcqWG6bGEdaqadaqaaiabigdaXiabgUcaRiabdogaJnaabmaabaGaeGymaeJaeyOeI0YaaeWaaeaacuWGHbqygaqeaiabgUcaRiabdggaHnaaBaaaleaacqaIYaGmcqWGibascqWGtbWuaeqaaaGccaGLOaGaayzkaaaacaGLOaGaayzkaaaacaGLOaGaayzkaaaacaGLOaGaayzkaaaaaa@4DEE@
Δu total (Δut)
a a t u ( − c + z ( 1 4 + c ( 1 − ( 3 a ¯ 4 + a 2 H S 2 ) ) ) ) MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacqWGHbqydaWgaaWcbaGaemyyaegabeaakiabdsha0jabdwha1naabmaabaGaeyOeI0Iaem4yamMaey4kaSIaemOEaO3aaeWaaeaadaWcaaqaaiabigdaXaqaaiabisda0aaacqGHRaWkcqWGJbWydaqadaqaaiabigdaXiabgkHiTmaabmaabaWaaSaaaeaacqaIZaWmcuWGHbqygaqeaaqaaiabisda0aaacqGHRaWkdaWcaaqaaiabdggaHnaaBaaaleaacqaIYaGmcqWGibascqWGtbWuaeqaaaGcbaGaeGOmaidaaaGaayjkaiaawMcaaaGaayjkaiaawMcaaaGaayjkaiaawMcaaaGaayjkaiaawMcaaaaa@4D1B@
a a t u ( − c + z ( 1 2 + c ( 1 − ( a ¯ 2 + a 2 H S ) ) ) ) MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacqWGHbqydaWgaaWcbaGaemyyaegabeaakiabdsha0jabdwha1naabmaabaGaeyOeI0Iaem4yamMaey4kaSIaemOEaO3aaeWaaeaadaWcaaqaaiabigdaXaqaaiabikdaYaaacqGHRaWkcqWGJbWydaqadaqaaiabigdaXiabgkHiTmaabmaabaWaaSaaaeaacuWGHbqygaqeaaqaaiabikdaYaaacqGHRaWkcqWGHbqydaWgaaWcbaGaeGOmaiJaemisaGKaem4uamfabeaaaOGaayjkaiaawMcaaaGaayjkaiaawMcaaaGaayjkaiaawMcaaaGaayjkaiaawMcaaaaa@4B1D@