From: The evolution of competition and policing: opposing selection within and among groups
Half sib
Full sib
Mean fitness ( w ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacuWG3bWDgaqeaaaa@2E3B@ )
1 - (z0 + 2qz a )
Δq within groups (Δqw)
p q z a ( p 2 z 1 H S + 2 p q + 1 2 z 2 H S + q 2 z 3 H S − 3 2 ) / 2 w ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=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@56D0@
p q z a ( p 2 z 2 F S + 2 p q z 4 F S + q 2 z 5 F S − 1 ) / 2 w ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacqWGWbaCcqWGXbqCcqWG6bGEdaWgaaWcbaGaemyyaegabeaakmaabmaabaWaaSaaaeaacqWGWbaCdaahaaWcbeqaaiabikdaYaaaaOqaaiabdQha6naaBaaaleaacqaIYaGmcqWGgbGrcqWGtbWuaeqaaaaakiabgUcaRmaalaaabaGaeGOmaiJaemiCaaNaemyCaehabaGaemOEaO3aaSbaaSqaaiabisda0iabdAeagjabdofatbqabaaaaOGaey4kaSYaaSaaaeaacqWGXbqCdaahaaWcbeqaaiabikdaYaaaaOqaaiabdQha6naaBaaaleaacqaI1aqncqWGgbGrcqWGtbWuaeqaaaaakiabgkHiTiabigdaXaGaayjkaiaawMcaaiabc+caViabikdaYiqbdEha3zaaraaaaa@52F0@
Δq between groups (Δqb)
-pqz a /4 w ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacuWG3bWDgaqeaaaa@2E3B@
-pqz a /2 w ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacuWG3bWDgaqeaaaa@2E3B@
Δq total (Δqt)
p q z a ( p 2 z 1 H S + 2 p q + 1 2 z 2 H S + q 2 z 3 H S − 2 ) / 2 w ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=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@55CA@
p q z a ( p 2 z 2 F S + 2 p q z 4 F S + q 2 z 5 F S − 2 ) / 2 w ¯ MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaacqWGWbaCcqWGXbqCcqWG6bGEdaWgaaWcbaGaemyyaegabeaakmaabmaabaWaaSaaaeaacqWGWbaCdaahaaWcbeqaaiabikdaYaaaaOqaaiabdQha6naaBaaaleaacqaIYaGmcqWGgbGrcqWGtbWuaeqaaaaakiabgUcaRmaalaaabaGaeGOmaiJaemiCaaNaemyCaehabaGaemOEaO3aaSbaaSqaaiabisda0iabdAeagjabdofatbqabaaaaOGaey4kaSYaaSaaaeaacqWGXbqCdaahaaWcbeqaaiabikdaYaaaaOqaaiabdQha6naaBaaaleaacqaI1aqncqWGgbGrcqWGtbWuaeqaaaaakiabgkHiTiabikdaYaGaayjkaiaawMcaaiabc+caViabikdaYiqbdEha3zaaraaaaa@52F2@